Distinguish current growth from coexistence under competition
Competition is a negative effect associated with shared limiting resources, but its strength can differ in each direction. Resource partitioning changes overlap, while predation, facilitation and environmental variation can modify outcomes. An ecological niche includes conditions and resource use; it is not simply a geographic address.
Learning outcome: Compare coefficients 0.5/0.5 with 1/1. Reconstruct current and rare-invasion rates, then explain why one census pair cannot identify long-run coexistence or a competition coefficient.
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Connect behavioral mechanisms, demographic stages, community interactions and ecosystem budgets while preserving denominators, causal alternatives and external fieldwork gates.
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At alpha = beta = 0.5, current population rates are 3.6 and 2.25 individuals per time. Rare-invasion rates are 0.12 and 0.075 per time, both positive. The current and invasion comparisons answer different questions.
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Can both species currently grow even if one cannot invade the other’s established population?
Experiment
Compare coefficients 0.5/0.5 with 1/1. Reconstruct current and rare-invasion rates, then explain why one census pair cannot identify long-run coexistence or a competition coefficient.
Compare current competition with invasion when rare
Worked teaching model · synthetic values · Continuous competition model dN1=.2N1[1−(N1+alpha N2)/100], dN2=.2N2[1−(N2+beta N1)/80], evaluated at N1=40,N2=30. Rare invasion into single-species equilibrium is distinct from current growth; no time integration or universal coexistence claim.
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Both rare-invasion rates are positive in this fixed competition model.
Current growth at one pair of abundances does not establish long-run coexistence. Resource partitioning can alter effective coefficients; facilitation, predation, spatial structure and changing resources are omitted.
| Quantity | Value | Unit |
|---|---|---|
| Current species-1 per-capita rate | 0.09000000 | per time |
| Current species-2 per-capita rate | 0.07500000 | per time |
| Current species-1 population rate | 3.600000 | individuals/time |
| Current species-2 population rate | 2.250000 | individuals/time |
| Species-1 rate when rare at species-2 equilibrium | 0.1200000 | per time |
| Species-2 rate when rare at species-1 equilibrium | 0.07500000 | per time |
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Both rare-invasion rates are positive in this fixed competition model.
Current growth at one pair of abundances does not establish long-run coexistence. Resource partitioning can alter effective coefficients; facilitation, predation, spatial structure and changing resources are omitted.
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At alpha = beta = 1, calculate current rates and both rare-invasion signs.
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