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SCIENCE · INVESTIGATE AND EXPLAIN

Explain richness turnover and island constraints

Use an immigration-extinction model without treating equilibrium as no ecological change.

Learning outcome: Use an immigration-extinction model without treating equilibrium as no ecological change.

Starting recap

Expected richness can remain constant while the species present change.

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Try

For c=0.3 and e=0.2 per year in a 100-species pool, expected equilibrium richness is 60.

Open the experiment

Predict

If c and e both double, does equilibrium richness change or only turnover and adjustment speed?

Experiment

Compare c/e ratios 0.2/0.1 and 0.4/0.2. Hold their ratio fixed; reconstruct the balanced rates. Then vary one coefficient alone.

Compare colonization and extinction pressures

Worked teaching model · synthetic values · Original expected-richness model: regional pool 100 species, immigration c(100−S), extinction eS. Equilibrium S*=100 c/(c+e), no speciation or habitat heterogeneity. Fractional expected richness is not a literal fraction of a counted species.

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Conditional calculations, not measured observations
QuantityValueUnit
Expected equilibrium richness60species
Immigration at equilibrium12species/yr
Extinction at equilibrium12species/yr
Adjustment time proxy2yr

Balanced rates can maintain expected richness while species identities turn over.

Isolation can reduce colonization; small area can increase extinction risk. Their mapping to coefficients is a hypothesis, not supplied measurement. A steady expected count does not mean zero arrivals or extinctions.

Predict before changing a control. Save two contrasting captures and explain what is held fixed. Displayed digits are calculation precision, not experimental uncertainty.

Balanced rates can maintain expected richness while species identities turn over.

Isolation can reduce colonization; small area can increase extinction risk. Their mapping to coefficients is a hypothesis, not supplied measurement. A steady expected count does not mean zero arrivals or extinctions.

Captures preserve recipes and replay their calculated results. They are synthetic model evidence, not physical measurements.

Understand

Area and isolation influence different processes

Island biogeography provides a framework for reasoning about colonization and local extinction. A site farther from a source pool may receive fewer colonists. A small habitat patch may support smaller populations and fewer habitat types, increasing some extinction risks. These are hypotheses about processes, not guarantees that every large or nearby island has a particular species count.

Habitat fragments on land can share some island-like features, but their surrounding matrix is not always an impermeable ocean. Species differ in dispersal and resource needs. Connectivity can facilitate movement, including movement of invasive species or pathogens, so a corridor proposal needs species-specific evidence.

Balance rates without freezing identities

Our original expected-richness model has a regional pool of 100 species. Immigration is c(100−S): fewer absent species remain to arrive as richness S increases. Extinction is eS: more represented species create more opportunities for local loss. At equilibrium, c(100−S)=eS, giving S*=100c/(c+e).

With c=0.3 and e=0.2 per year, expected richness is 60. Immigration is 0.3×40=12 species per year; extinction is 0.2×60=12. The expected count is stable while the identities can turn over. Fractional predictions in other settings are expectations, not a literal claim that half a species was observed.

Compare c=0.2,e=0.1 with c=0.4,e=0.2. Both give about 66.67 expected species. Balanced annual fluxes double from about 6.67 to 13.33. The adjustment-time proxy 1/(c+e) falls from about 3.33 to 1.67 years. The same equilibrium richness can conceal different turnover dynamics.

The model is not a measured island

The mapping of distance to c and area to e is not supplied by these controls. It must be estimated from observations or a more detailed model. Habitat quality, historical events, speciation, rescue effects and changing regional pools are omitted. The predicted equilibrium does not establish how rapidly every actual island reaches a stable state.

For a field comparison, select islands or fragments with measured areas and isolation, document survey effort, and consider habitat differences. A cross-sectional correlation between area and richness does not alone separate area from habitat quality or detection. Repeated species records can test turnover separately from a one-time count.

Transfer and solution tradeoffs

A restoration increases modeled colonization from 0.1 to 0.3 while extinction remains 0.2. Equilibrium richness changes from about 33.33 to 60. Explain the conditional prediction and ask which species are moving before claiming a conservation benefit. Greater total richness is not automatically greater persistence of a threatened native population.

A strong corridor recommendation specifies the target organisms, expected movement, possible unwanted transfers and monitoring needed to revise the design. It does not treat a balanced-rate equation as sufficient authorization for altering a habitat.

Colonization and evolved specialization are different processes

Colonization supplies arrivals; evolution changes inherited traits across generations. On islands, restricted resources and ecological opportunities can favor specialized use of food or territory. A broadly feeding introduced competitor can threaten an endemic specialist by consuming its limited resource, but nonnative status alone does not prove that interaction or impact.

An original island packet gives a native bird one usable seed type and a newcomer three usable types, including that seed. Predict whether the newcomer has an alternative during failure of the shared seed crop: yes, under the supplied diet information; the specialist does not. If the newcomer’s other foods also fail, its broad diet no longer guarantees an advantage. To support displacement rather than mere co-occurrence, compare shared-resource availability and native reproductive success with suitable reference observations. The equilibrium richness model counts turnover, not this evolutionary history or species-specific competition.

Source scope and evidence boundary

Original explanation, synthetic data and practice aligned with AP Environmental Science topic 2.3. Framework checked 30 October 2026: official topic map and Fall 2026 clarifications. The adaptation wording distinguishes inherited population changes over generations from individual need. Public numerical feedback is not independent assessment; virtual captures do not establish supervised fieldwork or actual ecosystem outcomes.

Exact-clause reconciliation checked 31 October 2026 using the official framework and Fall 2026 corrections. Added packets are original and observational or virtual only.

Apply

If c and e double together in S*=100 c/(c+e), equilibrium richness:

Public practice checks a specific claim. It does not establish independent mastery.

Use two contrasting captures, reconstruct the evidence, and state one changed-case prediction and one limitation. Use an immigration-extinction model without treating equilibrium as no ecological change.

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